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When macrophages are stimulated, for example by ligand interaction with the toll-like receptor 4 (TLR4), a signal transduction cascade is activated leading to enhanced expression of the ACOD1 gene

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GSH is known as a substrate in both conjugation reactions and reduction reactions, catalyzed by glutathione S-transferase enzymes in cytosol, microsomes, and mitochondria

It has potent effects of antioxidant, anti-inflammatory, anti-angiogenesis, and anti-cancer [131]